The Graf Lab @ The University of Alabama

email: dgraf(at) |
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dlgDr. Daniel L. Graf's
Curriculum Vitae

The following summarizes my career to-date. Use the following links to jump to various headings: Education, Professional Appointments, Grants, Awards & Fellowships, Service, Publications, Presentations & Posters, Other Scholarly Products, as well as student products.

My CV on this web page was last updated 30 August 2020.


1997-2001 Ph.D. Biology, University of Michigan, Ann Arbor, Michigan.
Dissertation: A Phylogenetic Perspective on the Evolution of the Unionoida (Mollusca Bivalvia Palaeoheterodonta): Using Pattern to Test Hypotheses of Macroevolutionary Process.
1994-1997 M.Sc. Biology, Northeastern University, Boston, Massachusetts.
Thesis: Morphology, Zoogeography, and Taxonomy of Fusconaia flava (Rafinesque, 1820) (Mollusca: Bivalvia: Unionidae) in the Upper Mississippi, Great Lakes, and Nelson River Basins.
1989-1993 B.Sc. Biology, College of Biological Sciences, University of Minnesota, St. Paul, Minnesota.

Professional Appointments

2018-present Associate Professor, Biology Department, University of Wisconsin-Stevens Point.
2012-2018 Assistant Professor, UWSP.
2008-present Research Associate, Academy of Natural Sciences of Philadelphia, Pennsylvania.
2004-present Research Associate, Illinois Natural History Survey, Champaign, Illinois.
2003-present Research Associate, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.
2008-2012 Assistant Professor and Full Member of the Graduate Faculty, Department of Biological Sciences, University of Alabama, Tuscaloosa, Alabama.
2002-2008 Assistant Curator, Malacology Department, Academy of Natural Sciences of Philadelphia, Pennsylvania.
2000-2001 Curatorial Assistant, Research Assistant and Collections Coordinator, Mollusk Division, University of Michigan Museum of Zoology, Ann Arbor, Michigan.
1995-1997 "Casual Help" and Curatorial Assistant, Malacology Department, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts.
1993-1994 Laboratory Assistant, Bell Museum of Natural History, University of Minnesota, St. Paul.

Recent Grants, Awards & Fellowships

2013-2014 ITIS, “Integrated Taxonomic Information System (ITIS) database development for freshwater mussels (Unionoida),” collaboration with Kevin Cummings (Illinois Natural History Survey): $13,000.
2013-2014 Undergraduate Education Initiative (UEI), UWSP, “Curation of the Daniel Bereza Collection of Freshwater Mussels at the National Museum of Natural History”: $3880.
2007-2013 National Science Foundation, "Phylogeny on the half-shell — Assembling the Bivalve Tree of Life." Senior Investigator with Gonzalo Giribet (Museum of Comparative Zoology), DEB-AToL-0732903, $137,574.
2007-2008 IUCN Freshwater Biodiversity Unit, Central Africa freshwater biodiversity assessment project, "Unionoida Assessment." 510 euros.
2006-2009 National Science Foundation, "Evolution in a Vortex: An Inventory of the Fishes and Mollusks of the Lower Congo River Rapids." Collaboration with Melanie Stiassny (American Museum of Natural History), DEB-BSI-0542575: $139,550 (including a supplemental award of $14,295).
2003-2008 National Science Foundation, "Systematic Revision of the Etherioidea, the Freshwater Mussels (Mollusca: Unionoida) of the Gondwanan Continents." Collaboration with Kevin Cummings (Illinois Natural History Survey), DEB-REVSYS-0316125: $199,805.

Recent Service

2007-present Journal of Molluscan Studies, Associate Editor.
2009-present IUCN SCC Mollusca Specialist Group, member.
2011-present Freshwater Mollusk Biology and Conservation, editorial board.
2009-2018 Malacologia, editorial board.
2015-2017 American Malacological Society, Council member at-large.
2002-present NSF Proposal Reviews.
2000-present Numerous journal (and other scholarly publications) reviews.

Recent Publications

† indicates student collaborators. Click here for a complete list of my publications.

Pfeiffer, J.M., D.L. Graf, K.S. Cummings & L.M. Page. 2021. Taxonomic revision of a radiation of Southeast Asian freshwater mussels (Unionidae: Gonideinae: Contradentini + Rectidentini). Invertebrate Systematics 35: 394-470.

Graf, D.L. & K.S. Cummings. 2021. A 'big data' approach to global freshwater mussel diversity (Bivalvia: Unionoida), with an updated checklist of genera and species. Journal of Molluscan Studies 87(1): eyaa034, 36 pp.

Böhm, M., N.I. Dewhurst-Richman, M. Seddon, S.E.H. Ledger, C. Albrecht, D.Allen, A.E. Bogan, J. Cordeiro, K.S. Cummings, A. Cuttelod, G. Darrigran, W. Darwall, Z. Fehér, C. Gibson, D.L. Graf, F. Köhler, M. Lopes-Lima, G. Pastorino, K.E. Perez, K. Smith, D. van Damme, M.V. Vinarski, T. von Proschwitz, T. von Rintelen, D.C. Aldridge, N.A. Aravind, P.B. Budha, C. Clavijo, D. Van Tu, O. Gargominy, M. Ghamizi, M. Haase, C. Hilton-Taylor, P.D. Johnson, Ü. Kebapçı, J. Lajtner, C.N. Lange, D.A.W. Lepitzki, A. Martínez-Ortí, E.A. Moorkens, E. Neubert, C.M. Pollock, V. Prié, C. Radea, R. Ramirez, M.A. Ramos, S.B. Santos, R. Slapnik, M.O. Son, A.-S. Stensgaard, & B. Collen. 2021. The conservation status of the world’s freshwater molluscs. Hydrobiologia 848: 3231–3254.

Graf, D.L. & K.S. Cummings. 2019. [Six chapters] Unionidae, Margaritiferidae, Hyriidae, Etheriidae, Iridinidae, Mycetopodidae. In C. Lydeard & K.S. Cummings (eds.). Freshwater Mollusks of the World: A Distribution Atlas. Johns Hopkins University Press, Baltimore, MD.

† Pfeiffer, J.M., D.L. Graf, K.S. Cummings & L.M. Page. 2018. Molecular phylogeny and taxonomic revision of two enigmatic freshwater mussel genera (Bivalvia: Unionidae incertae sedis: Harmandia and Unionetta) reveals a diverse clade of southeast Asian Parreysiinae. Journal of Molluscan Studies 84: 404-416.

Recent Presentations & Posters

† indicates student collaborators. Click here for a complete list of presentations & posters.

Graf, D.L. 2021. Good, Clean Family Fun: Biodiversity informatics of freshwater bivalves above the species level [oral presentation]. American Malacological Society, 14-18 June 2021, virtual meeting.

Graf, D.L. 2020. A Biodiversity Informatics Approach to Family-group Level Classification of Freshwater Mussels (Bivalvia: Unionoida) [oral presentation]. American Malacological Society, 13-14 July 2020, virtual meeting.

Graf, D.L. 2019. A big data approach to biodiversity research. Great Lakes Analytics Conference, 4 October 2019, University of Wisconsin-Stevens Point, Stevens Point, WI.

Cummings, K.S., C.R. Randklev, K. Inoue, J.S. Tiemann, N.A. Johnson, J.M. Pfeiffer, D.L. Graf, T. Miller, A.L. Burden & E. Naranjo-Garcia. 2019. The Freshwater Mussels of México (Unionidae & Mycetopodidae) [oral presentation]. Biennial Freshwater Mollusk Conservation Society Symposium, 14-18 April 2019, San Antonio, TX.        

Cummings, K.S., D.L. Graf, C. Randklev, K. Inoue, A. Pieri & J.S. Tiemann. 2018. The freshwater mussels of México [oral presentations]. American Malacological Society, 19-23 June 2018, Honolulu, HI.

Graf, D.L. 2017. Conflict Clade Analysis (CCA) of the freshwater mussel subfamily Unioninae (Bivalvia: Unionoida) [oral presentation]. American Malacological Society, 17-21 July 2017, University of Delaware, Newark, DE.

Other Recent Scholarly Products

Graf, D.L. & K.S. Cummings. 2002-2023. "The MUSSEL Project Web Site." Web site dedicated to explaining freshwater mussel diversity and evolution:

Graf, D.L. 2012-2023. "WInvertebrates: Towards an Account of the Freshwater Invertebrate taxa of Wisconsin, USA." Web site dedicated to synthesizing information of the diversity and zoology of the non-parasite, non-insect invertebrates of Wisconsin and adjacent areas:

Graf, D.L. 2008. Book review of "Freshwater Mussel Ecology- A Multifactor Approach to Distribution and Abundance by David L. Strayer." The Malaclogist 51:!musselreview51.htm

Recent Student Work

Dunn, I.R., L.C. Trittelwitz, J.L. Bielak, A.S. Mehus, & M.M. Tokarski. 2020. Freshwater mussel abundance and species richness with varying substrate [poster]. 2020 Jim & Katie Krause CNR Student Research Symposium, UWSP.

Bielak, J.L., A.S. Mehus, M.M. Tokarski, L.C. Trittelwitz & I.R. Dunn. 2020. Freshwater mussel species richness and population patterns in Portage County [poster]. 2020 Jim & Katie Krause CNR Student Research Symposium, UWSP.

Roeske-Kretzer, O. & L. Taylor. 2019. Biogeography of freshwater mussels on the Wisconsin Natural Heritage Working List [poster]. UWSP College of Letters and Science Undergraduate Research Symposium, 3 May 2019.

Dykstra, R. 2018. Updated assessment of the biodiversity of freshwater mussels in Minnesota, Wisconsin, and Michigan [poster]. UWSP College of Letters and Science Undergraduate Research Symposium, 4 May 2018.

Hausmann, E. 2017. Phase I: Mesoamerican freshwater mussels (Bivalvia: Unionidae) [poster]. UWSP College of Letters and Science Undergraduate Research Symposium, 5 May 2017.

Last updated 23 June 2023